Wydarzenia przedstawione w grze stanowią bezpośrednią kontynuację wątków znanych z Assassin's Creed II. Gracz ponownie wciela się w postać Ezio Auditore de Firenze, który trudni się rzemiosłem tytułowego Assassina. Tym razem cała intryga rozgrywa się w murach wiecznego miasta, zaś koncentruje się na brutalnym konflikcie pomiędzy tajemniczym zgromadzeniem zabójców a zakonem Templariuszy.

Assassin's Creed: Brotherhood kontynuuje wszystko to, co widzieliśmy w częściach poprzednich oraz dodaje od siebie garść świeżych pomysłów. Sedno zabawy ponownie stanowi eksploracja miasta, wykonywanie różnorakich zleceń oraz widowiskowe ucieczki. W trakcie przygody Ezio spotka na swojej drodze kilka historycznych postaci, m.in. Leonarda da Vinci oraz Niccolo Machiavelliego.

W stosunku do poprzednich odsłon cyklu Assassin's Creed, rozbudowuje zestaw dostepnych narzędzi. Wśród nowych zabawek znajdziemy chociażby wachlarz, strzykawkę z trucizną, a nawet prototypowy spadochron. Co ciekawe, w sytuacjach podbramkowych będziemy mogli skorzystać z pomocnej dłoni innych Assassinów, którzy przebywają w mieście.

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